Ecstatic Bees not retaining traits

[Former user]

I have an ecstatic queen in my apiary, i have modified it with beatific effect, flower pollination, longest life, fast production, and maximum fertility. However as soon as the queen dies out, the drones and princess do not retain any of these traits, and they all become the inactive trait again. And yes i did modify the first Queen's inactive and active traits. My other bees are doing fine, i have a diamond queen, a couple emerald queens, an imperial and an industrious and they all retain their traits.

[Alex Binnie]

This is how the breeding mechanics work, you need to inoculate the princess, not the queen